Almost all organisms use carbohydrates as sources of energy. In addition, some carbohydrates serve as structural materials. Carbohydrates are molecules composed of carbon, hydrogen, and oxygen; the ratio of hydrogen atoms to oxygen atoms is 2:1.
Simple carbohydrates, commonly referred to as sugars, can be monosaccharides if they are composed of single molecules, or disaccharides if they are composed of two molecules. The most important monosaccharide is glucose, a carbohydrate with the molecular formula C6H12O6. Glucose is the basic form of fuel in living things. It is soluble and is transported by body fluids to all cells, where it is metabolized to release its energy. Glucose is the starting material for cellular respiration, and it is the main product of photosynthesis.
1 shows that in the synthesis of sucrose, a water molecule is produced. The process is therefore called a dehydration. The reversal of the process is hydrolysis, a process in which the molecule is split and the elements of water are added.) Lactose is composed of glucose and galactose units.
Glucose and fructose molecules combine to form the disaccharide sucrose.
Complex carbohydrates are known as polysaccharides. Polysaccharides are formed by linking innumerable monosaccharides. Among the most important polysaccharides are the starches, which are composed of hundreds or thousands of glucose units linked to one another. Starches serve as a storage form for carbohydrates. Much of the world's human population satisfies its energy needs with the starches of rice, wheat, corn, and potatoes.
Two other important polysaccharides are glycogen and cellulose. Glycogen is also composed of thousands of glucose units, but the units are bonded in a different pattern than in starches. Glycogen is the form in which glucose is stored in the human liver. Cellulose is used primarily as a structural carbohydrate. It is also composed of glucose units, but the units cannot be released from one another except by a few species of organisms. Wood is composed chiefly of cellulose, as are plant cell walls. Cotton fabric and paper are commercial cellulose products.
Lipids are organic molecules composed of carbon, hydrogen, and oxygen atoms. The ratio of hydrogen atoms to oxygen atoms is much higher in lipids than in carbohydrates. Lipids include steroids (the material of which many hormones are composed), waxes, and fats.
2 ). A glycerol molecule contains three hydroxyl (—OH) groups. A fatty acid is a long chain of carbon atoms (from 4 to 24) with a carboxyl (—COOH) group at one end. The fatty acids in a fat may be all alike or they may all be different. They are bound to the glycerol molecule by a process that involves the removal of water.
A fat molecule is constructed by combining a glycerol molecule with three fatty acid molecules. (Two saturated and one unsaturated fatty acids are shown for comparison.) The constructed molecule is at the bottom.
Certain fatty acids have one or more double bonds in their molecules. Fats that include these molecules are unsaturated fats. Other fatty acids have no double bonds. Fats that include these fatty acids are saturated fats. In most human health situations, the consumption of unsaturated fats is preferred to the consumption of saturated fats.
Fats stored in cells usually form clear oil droplets called globules because fats do not dissolve in water. Plants often store fats in their seeds, and animals store fats in large, clear globules in the cells of adipose tissue. The fats in adipose tissue contain much concentrated energy. Hence, they serve as a reserve energy supply to the organism. The enzyme lipase breaks down fats into fatty acids and glycerol in the human digestive system.
3 ), which contain carbon, hydrogen, oxygen, and nitrogen atoms. Certain amino acids also have sulfur atoms, phosphorous, or other trace elements such as iron or copper.
The structure and chemistry of amino acids. When two amino acids are joined in a dipeptide, the —OH of one amino acid is removed, and the —H of the second is removed. A dipeptide bond (right) forms to join the amino acids together.
3 ) has an amino (—NH2) group, a carboxyl (—COOH) group, and a group of atoms called an —R group (where R stands for radical). The amino acids differ depending on the nature of the —R group, as shown in the middle illustration of Figure 3 . Examples of amino acids are alanine, valine, glutamic acid, tryptophan, tyrosine, and histidine.
The removal of water molecules links amino acids to form a protein. The process is called dehydration synthesis, and a byproduct of the synthesis is water. The links forged between the amino acids are peptide bonds, and small proteins are often called peptides.
All living things depend on proteins for their existence. Proteins are the major molecules from which living things are constructed. Certain proteins are dissolved or suspended in the watery substance of the cells, while others are incorporated into various structures of the cells. Proteins are also found as supporting and strengthening materials in tissues outside of cells. Bone, cartilage, tendons, and ligaments are all composed of protein.
One essential use of proteins is in the construction of enzymes. Enzymes catalyze the chemical reactions that take place within cells. They are not used up in a reaction; rather, they remain available to catalyze succeeding reactions.
Every species manufactures proteins unique to that species. The information for synthesizing the unique proteins is located in the nucleus of the cell. The so-calledgenetic code specifies the amino acid sequence in proteins. Hence, the genetic code regulates the chemistry taking place within a cell. Proteins also can serve as a reserve source of energy for the cell. When the amino group is removed from an amino acid, the resulting compound is energy rich.
Like proteins, nucleic acids are very large molecules. The nucleic acids are composed of smaller units called nucleotides. Each nucleotide contains a carbohydrate molecule, a phosphate group, and a nitrogen-containing molecule that because of its properties is a nitrogenous base.
Living organisms have two important nucleic acids. One type is deoxyribonucleic acid, or DNA. The other is ribonucleic acid, or RNA. DNA is found primarily in the nucleus of the cell, while RNA is found in both the nucleus and the cytoplasm, a semi-liquid substance that composes the foundation of the cell.
DNA and RNA differ from one another in their components. DNA contains the carbohydrate deoxyribose, while RNA has ribose. In addition, DNA contains the base thymine, while RNA has uracil.
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